A change of allegiance leads to the arrest of Andréas. Despite their orders, Caplan and Morlighem continue to investigate. They manage to dismantle a drug ring involving Russians and Turks. Roxane and Hélène have fallen into the hands of Vogel, who gives Caplan a grotesque choice: which one does he want him to kill? While the various Vor clans kill each other in Paris, the lives of Caplan, Morlighem and Roxane are about to take a dramatic new turn.

Oriane has managed to work her way into Caplan’s group as an interpreter. She is bent on vengeance. Caplan has his gun in his hand when the Russians launch an assault on their stakeout. Flashback: five years earlier. A war within police ranks between Rossi/Caplan’s unit and Vibert’s. During an operation, Vibert shoots a suspect in cold blood. Turned in by Caplan, Vibert is sent to prison. Oriane, who is in fact Vibert’s daughter, has been seeking to avenge her father ever since.

After his car blew up, Theo was taken to hospital in a critical state. Caplan and Morlighem go to Vogel's place, determined to avenge their companion. Vogel finds an ally in the shape of Orianne Beridzé, who also wants to eliminate Caplan. Her plan is to force her father, an ex-cop, to contact Caplan and put him on the tail of a trafficking network run by the Russian Mafia. Meanwhile, Roxane is deeply upset about Theo and watches over him day and night.

With the death of Vissarionovitch – who ruled as 'boss of bosses' over several Russian-speaking mafia clans – a bloody war of succession is shaping up between the two pretenders: Jordania and Pavlovitch. Roxane continues to operate alone, tracking down Vogel’s sister thanks to information provided by Vanderbeke. In the meantime, Caplan, Morlighem and Megrelishvili stake out the funeral where Vissiniarovitch’s burial service is due to take place. Odessa, a female contract killer, sets up shop in a hotel opposite, ready to strike.

After an assassination attempt, Jordania is placed under police surveillance in hospital. His adversary, Pavlovitch, teams up with Atom Parajdanov. They are trying to get to Jordania any way they can. Caplan offers him a deal: hand over information on his rivals (become a 'stoukatch' – a traitor) in exchange for police protection. Meanwhile, Vogel and Oriane take Hélène Rossi hostage to put pressure on Caplan.

The Russians got the better of Jordania, who was killed in the course of the attack. But how did the maximum security safe house get located so quickly? Caplan’s suspicions fall on Andréas. Under questioning, Andréas explains that he is working to a plan drawn up by Georgian police in agreement with their French counterparts, the aim of which is to lure the top boss of the Vor V Zakone from Moscow and to arrest him in France. And Caplan’s unit had better keep out of their way.

The Pavlovitch clan organizes the kidnapping of Salomé and her son to force Jordania into the open. Jordania has no option but to accept Caplan’s deal and turn traitor to protect his family. Roxane meanwhile has tracked down Vogel again. He is at her mercy, but she can’t bring herself to pull the trigger. She rejoins her unit, assigned as bodyguard for Jordania, and they head to a safe house 30 miles outside of Paris.

Caplan and Morlighem have intercepted a truck from a company belonging to the Russians and uncover a major prostitution ring. They are aided by a cop from Georgia, Andreas Megrelishvili, who informs them that a war between clans of the Vor V Zakone Mafia from different post-Soviet states is about to break out in Paris. Meanwhile, Roxane, who wants revenge for Vogel, strikes a deal with the former prosecutor, Vanderbeke.