Annabelle Neilson is back on the scene after a horrible accident last season while Julie Montagu struggles with being a working mom now that her husband is busy running their family’s Mapperton estate. Marissa Hermer sets out to throw a lavish Thanksgiving dinner for her society friends, but as the holiday approaches, Juliet Angus and Caroline Stanbury forge an alliance that could spell the end of Juliet’s friendship with Marissa.

Caroline Fleming organizes an eccentric holiday party. Annabelle tries to help Julie improve her confidence. Caroline Stanbury attempts to deal with the problems of running a business.

Much to Julie's chagrin, Caroline Stanbury instructs her on the emerging health food industry; the women celebrate New Year's Eve in the penthouse of a London hotel, where Caroline's antics threaten to push Juliet and Julie over the edge.

The ladies deal with fallout from naughty New Year's Eve high jinks. Meanwhile, Marissa receives positive press for her restaurant; Caroline Fleming visits her grandmother in Denmark; and Caroline Stanbury tries to evade gossip.

Julie extends an olive branch to Caroline Stanbury in spite of their fallout on New Year's Eve; Marissa seeks Annabelle's advice in anticipation of the launch of her new restaurant; Julie looks for funding for her snack-food startup.

After being on the outs with the ladies for weeks, Juliet attempts to turn a new leaf when she invites everyone out for a night of bowling -- but whether she can convince them she’s changed remains to be seen. Caroline Stanbury faces the unsettling reality of having to scale down her staff in order to appease investors and keep her business afloat. Julie takes Annabelle and Caroline Fleming to her uncle’s Beaulieu estate to generate ideas for saving Mapperton.

Marissa must work to salvage her friendship with Caroline Fleming after insulting her for dating a younger man. Julie once again fulfills her reputation as "Loose Lips" Montagu when she hosts a shooting weekend for the girls. Caroline Stanbury struggles to remain emotionally stable while her Gift Library continues its downward spiral.

Marissa and Juliet come back together to watch Annabelle take to the catwalk with Naomi Campbell. Caroline Stanbury reaches a tipping point with Gift Library. Caroline Fleming invites the ladies to her homeland of Demark where they are shocked by her celebrity. Though they are far from home, the mounting tension between the ladies cannot be contained, as Annabelle voices her opinions - loudly.

The tension bubbling between Annabelle and Caroline Stanbury boils over at Caroline Fleming’s enormous castle. The baroness visits her mother’s grave and faces her father for the first time since Marissa’s meddling. The ladies uncover the reason behind Annabelle’s attitude.

In the season finale, Caroline Stanbury adjusts to life without Gift Library. The ladies wonder if Annabelle will trust them enough to show at their final winter bash, where a new Queen Bee is buzzing. Each of the Ladies’ future is gleaming.