Just as it seems lion cubs Prince and Ziggy have finally been accepted into the pride a jealous lioness intends to get rid of them. Rogue males invade the territory of the Hollywood pride threatening their cubs.

Young cub Pip's adventurous nature soon gets him into trouble as he wanders away from the pride. A lost, lonely young lion is easy prey for prowling leopards and hyenas. Meanwhile Pip's cousins Ziggy and Prince have troubles of their own. They've survived an attack from a rival male lion, but now their mum Rosa is trying to re-join her pride. Ziggy and Prince's new step-dads are far from happy.

Tensions rise between two lion prides as their territories overlap - which has dangerous consequences for an orphaned elephant searching for its family group. Two dominant males try to regain control of their old pride.