They're soul mates who've never met, a man and woman with parallel lives for whom first love is long gone and perfect lovers are yet to appear. Gil is an American comedy writer who came to the English countryside for what he believed was true love, while Alice is a perfume account manager for a London department store, set to move from the country and her pricey Brighton flat because of the commute. Milly and Cleo are forever trying to sniff out Mr. right for Alice, but can they orchestrate a meeting?

Alice's birthday brings presents and dates she didn't ask for, and Gil somehow gets more involved in Irene's life than he'd like.

Alice's part in a company presentation offends her, but can she stand up to a superior whose review could determine her future at the store? Meanwhile, Gil tries a blind date with a fan, but is distracted by concerns for Irene and her date.

Alice and Gil follow different dating paths; she is set up and meets an artist, and he is set up but continually stood up, by the same actress.

Infatuations preoccupy both Alice and Gil... For very different reasons.

While helping out a cousin, Alice believes she may have met her soul mate. Elsewhere, when Gil's fantasy comes true, he's a bit disconcerted.

Alice worries that her manager may be after more than a business meeting when she's summoned to the woman's hotel room, Cleo obsesses over a bad review and Milly is strangely drawn to a man's shadow.

Alice bumps into what may be the "Perfect Man" at a cash machine, but their first date includes meeting the family and a spectre from his past. Elsewhere, Cleo and Milly devise a test for Darren's love of Milly.

Alice's yearly lunch with a college chum is interrupted by an obsessed student; Milly and Cleo try out the speed dating fad.

Alice's encounter with the anti-terrorism unit of the local police may have something to do with the romantic deceit she engaged in with her latest beau, and Cleo indulges in some sexual games of her own.

Alice is swept into the posh life of Matthew Chater; Milly and Cleo find themselves arguing over a tee-shirt, a man, and a funeral.

Cleo and Milly get in trouble when they access personal health records at the store's chemist in an effort to screen potential dates for Alice; meanwhile, Alice's boyfriend's job may be its own deterrent.

Alice's finances force her to finally let her Brighton flat and look for something cheaper in London.

Everything seems to be going well with Alice's new bed-in, Douglas, but a message from his ex-girlfriend will plunge them into another round of humiliations and disappointment.

Alice is excited by her opportunity to see a play directed by a comedy actor she admires.

Alice's life is in turmoil, although her housing situation has stabilised, and she wonders if her choice about Douglas was correct.

Getting used to a new flatmate is difficult, unfortunately Alice's date provides no distraction as it descends into the drama that's typical of her outings.

Alice's loneliness becomes pervasive. Is the solution another date, or will the grand plan finally be revealed?