Winter has finally come to a close, spelling big changes for Tom, who bids farewell to an old friend and packs his bags for Florida. In the Ruby Valley, Rich grooms his youngest hound to take over the pack, but an influx of cougars finds the pup in over his head. In North Carolina, Eustace takes a dangerous trip down a rapid river to deliver the canoe he built to pay off his loan. In Maine, Charlie takes his furs to market and hopes to break even on his rookie season, while in Alaska, Marty enjoys a reunion with his family, then makes big plans for the future.

Marty rushes to make the most of an extra pair of hands. Tom's grandson comes for a visit but soon learns that there's no such thing as vacation in the Yaak. Charlie heads out on a hunt for shed moose antlers. And in Montana's Ruby Valley, Rich Lewis' dogs are hot on the scent of a mountain lion when their hunt takes a dangerous turn.

Kidd and Harry traverse through intense whitewater to reach the spawning grounds of steelhead trout. Jake loses his dogs after they follow lion tracks into remote and unfamiliar territory. Paul checks his traps and takes home the biggest prize on the mountain. Eustace mills massive beams to repair a deteriorating dock at a local marina. Aron and Jen mill all their own lumber and rebuild a cabin roof.

In Alaska, after spending the night under the stars in subzero conditions Marty struggles to get his snow machine out of a stubborn overflow. In the Ruby Valley, Rich takes his oldest dog, Brandy and youngest dog, Hatchet, out on a special training mission. Eustace and Preston set off on a crucial deer hunt, but are interrupted by loud intruders. In the Yaak Valley, Tom is called by his friend, Tim to help with a beaver problem and takes the opportunity to teach an important lesson; and in Maine, Charlie and Jim head to the frozen river to carve out 100 pound ice chunks for Charlie's root cellar.

Tom and Sean brave the bitter cold in search of better trapping grounds; Jake seeks revenge on the lion that injured his hound; Eustace and Raleigh struggle to fix a storm-damaged bridge that connects Turtle Island to the world outside; Mary forges her first Ozark blade; and Josh loses a bison to a pack of hungry wolves.

Jason's bladesmithing business suffers a critical setback when his hand is badly cut on the job; Jake's latest lion hunt turns into a rescue mission when he discovers his dogs are on a collision course with a pack of wolves; Kidd and Harry break a colt for a backcountry client; and Josh salvages all he can from a bison that was killed by wolves.

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In the Yaak Valley, Tom Oar teams up with his brother Jack to help return Jack's prized hunting falcon back to the wild. In southern Montana, Rich Lewis and his wife Diane need to move their cattle herd, but will the predators get to them first? Marty starts wrapping up a successful season, but the elements won't let him leave Alaska without a fight. Eustace and Preston realize they have to contend with coyotes who are putting their wild game at risk. And in Maine, rookie Charlie Tucker struggles to re-start his season after an injury left him with one working arm.

In the Season 4 premiere, unpredictable extreme conditions lead to the survivalists finding new ways to adapt.

Marty makes a gamble that changes everything; a storm batters Eustace and Preston during their final push.

Mike's bear scout turns into an alpine goat hunt; Jake brings in the big dogs to settle an old score with a problem lion; Tom and Sean check beaver sets to see if their new trapping grounds will pay off; Jason leans on the family to overcome a looming deadline and a broken hand; and Kidd and Harry run a backcountry gauntlet to deliver a young horse to a remote buyer.

Josh witnesses a bison give birth to the first calf of spring; Kidd and Harry load a mule into their boat and transport it downriver through raging rapids; Jake runs two mountain lions up the same tree; Eustace receives help from his neighbors as they rush to repair the roof of a damaged hay barn; Mike adds off-grid utilities to his remote hunting cabin.

Jake and the pups make a grisly discovery in new territory; Josh rebuilds an antique windmill to power his outpost cabin; Kidd and Harry brave raging rapids to retrieve a capsized canoe; Martha and Elli clear the road of fallen trees.

Eustace hunts for his winter meat in the pre-dawn hours. Jake harvests a deer with bow and arrow. Tom helps his friend Paul prepare for trapping season. Ray stalks elk through knee deep powder, and spends a frigid night on the mountain. Aron hauls his winter supplies over a treacherous off-road trail, but the final stretch of the mission proves most daunting.

Tom packs a traditional muzzleloader on a winter deer hunt with Sean and Hank; Jake and Anika complete a successful elk hunt with a backbreaking pack out; Aron and Jen cut down dead trees that threaten to destroy their cabin.

Eustace and his neighbor train a stubborn stallion; Jake and his team of dogs successfully run off their first mountain lion of the season; Paul tracks and kills a young buck; Aron hunts for moose, while Jen fishes for rainbow trout.

Jake and his dogs tree three cats in one day; Paul prepares his haul of winter fur for market and explores a new piece of land to call his own.

Marty Meierotto arrives at his Alaska trapline, but the unseasonably warm weather is impeding his plans. In North Carolina, Eustace Conway discovers two headless chickens in his henhouse, and begins a hunt for the predator responsible. In northwest Montana, Tom Oar discovers a grizzly bear has been prowling his property, while in the southern corner of the state, Rich Lewis frantically searches for his missing hound, Turbo.

Desperate for food after a grizzly bear ravaged his garden, Tom Oar embarks on a last-ditch deer hunt. Marty Meierotto begins to establish his winter trapline, breaking fresh trail in the uncharted wild. In the Ruby Valley of Montana, Rich Lewis pursues a marauding wolf pack that has slaughtered a neighbor's cattle. And in the Blue Ridge Mountains of North Carolina, Eustace Conway must deal with Scout, a careless intern with "a dose of city boy in him.

After a successful deer hunt, Tom rushes to prepare the meat before it attracts hungry grizzly bears. In North Carolina, Eustace takes on a grueling job that he needs to complete in order to make his first loan payment. Marty's mechanical problems return and leave him stranded in the cold Alaskan night. In Montana's Ruby Valley, Rich postpones an elk hunt when he discovers evidence of a lion poacher. Rookie mountain man Charlie Tucker begins his trapping season in Maine's Great North Woods.

Rich and his hounds track Rich's nemesis, the killer mountain lion Three-Toes, across the length of Montana's Ruby Valley. In Alaska, a young prot g brings Marty the new snowmobile engine he's been waiting for. Eustace races the winter frost to seed his field in North Carolina and in Northwest Montana, Tom receives a commission from a neighbor to build a primitive bow, with a deadline that puts his endurance to the test.

Logger turned Mountain Man, Charlie Tucker's luck takes a turn for the worse in the sub-zero North Maine Woods as a rookie mistake nearly costs him everything. In Alaska, Marty and Andy split up to cover more ground with the plan to rendezvous at dusk. But as night falls, Marty is missing in action. In North Carolina, Eustace puts his life on the line to help a neighbor in need, and in Montana's Ruby Valley, Rich Lewis sets off after a bobcat which has been eating his livestock, and the cat draws first blood, injuring two of his dogs before Rich vows revenge.

Rich's cougar hunt takes a dark turn in Montana. Marty competes with a wolf pack for food. Tom and Will race an oncoming storm that puts them in jeopardy. And in Maine, Charlie suspects sabotage.

Charlie spends the night injured and alone in Maine. In Montana Tom investigates signs of a predator lurking too close to home. Eustace rushes to raise money to make his loan payment. And in Idaho, new mountain man George Michaud tries to make a living running his dog sled through the peaks of the mighty Tetons.

Tom struggles to keep a deadly wolf pack off his land despite an injury. Rich tracks a large buck over tough terrain on his mid-winter hunting trip. Eustace makes some much-needed repairs and George is forced to make his way across the Tetons on foot across before bad weather hits.