After defeating Kunzite, regaining their past memories, and discovering the Dark Kingdom's hideout the Sailor Senshi head for the enemy, but they must prepare for the ultimate sacrifice.

Mamoru is forced to confront Zoisite in a duel over the Rainbow Crystals, but Usagi inadvertently becomes involved.

With her friends killed, Sailor Moon is transported inside the Dark Kingdom to face Queen Beryl and the brainwashed Prince Endymion.

Still in love with Nephrite, Naru becomes an unwilling tool in the power struggle between Nephrite and Zoisite.

Queen Beryl partially awakens the evil entity Metalia and sends Zoisite and Kunzite to find the Seven Great Youma and their Rainbow Crystals. Meanwhile, the tomboyish Makoto Kino transfers into Usagi's school and falls for a crane-game whiz at the arcade.

Usagi and Mamoru remember their former lives as Princess Serenity and Prince Endymion. The Dark Kingdom kidnaps the injured Mamoru, but Queen Beryl executes Zoisite for his attempt to kill him. Luna and Artemis tell the Sailor Guardians about their tragic past.

A blast of energy sends the Guardians to the ruined Moon Kingdom, where they meet a hologram of Queen Serenity and learn about their past. Luna and Artemis find the entrance to the Dark Kingdom. Kunzite attacks, but is finally defeated by Sailor Moon.

The Sailor Guardians go to the Dream Land amusement park, where visitors have reportedly gone missing. The team splits into two groups: Usagi and Rei and Ami and Luna. But, much to Rei's frustration, Usagi is more focused on playing than finding the missing people.

Zoisite disguises himself as Sailor Moon to lure Tuxedo Mask and the Sailor Guardians out into the open. The Dark Kingdom injures Tuxedo Mask's identity, but they are foiled by the appearance of Sailor Venus.

Zoisite decides to use Nephrite's recently discovered romantic feelings for Naru to manipulate him into handing over the Black Crystal to him.

Several people who take the bus near the Hikawa Shrine have gone missing. Usagi, Ami, and Luna decide to investigate by visiting the shrine. There, they meet Rei Hino, a shrine maiden with strong spiritual powers.

Queen Beryl is increasingly furious with Jadeite for his repeated failures; he starts up a theme park, but the general public notices the large number of disappearances. Usagi and Rei, who is now Sailor Mars, begin their combative friendship.

Usagi Tsukino is a 14-year-old schoolgirl who is very cheerful, but clumsy and is always finding herself into unwanted trouble. One day, Usagi meets a talking black cat named Luna, who reveals that dark forces are threatening the lives of the innocent and that Usagi has the power to stop them. Using a magical brooch, Usagi is transformed into Sailor Moon, the chosen guardian who fights for love and justice.

Luna discovers that she has an admirer, in the form of an overweight tomcat named Rhett Butler. Meanwhile, the Sailor Guardians and Zoisite compete for the last Rainbow Crystal.

Usagi befriends Yusuke Amade, a jazz musician who is being followed by a Youma, and helps him confess to the woman he loves.

Both Luna and Queen Beryl wonder if a visiting princess could be the one the Sailor Senshi has been looking for. Mamoru dreams he is Tuxedo Mask and a girl is asking him for the Silver Crystal. At the ball, Usagi gets her first kiss.

Usagi and Mamoru are asked to model for the reclusive painter Yumemi Yumeno, who is unexpectedly targeted by Zoisite.

Makoto, like Usagi, develops a crush on Motoki. The two then decide to pursue him simultaneously, even after finding out that he has a girlfriend, who, meanwhile, is targeted by Zoisite and Kunzite.

One of Queen Beryl's youma offers to help Jadeite with his wicked plans. Rei wins tickets on a luxury cruise ship and, since she has no boyfriend, asks Ami to accompany her. A jealous Usagi sneaks aboard after them.

Usagi continues to be attracted to both Motoki and Tuxedo Mask. Threatened with Eternal Sleep if should he fail again, Jadeite confronts the Sailor Guardians at an airport.

Ami meets Ryo Urawa, a clairvoyant boy who has a crush on her. However, Urawa's power makes him aware that he will soon be transformed into a Youma by Zoisite.

Minako tries to cheer Usagi up by taking her to a hair salon, leading to the discovery that Tuxedo Mask is still alive, but now very different.

Usagi and her friends visit the Moonlight Course skiing resort, unaware that Kunzite has sabotaged the Moon Princess Contest in order to unmask Sailor Moon.

Queen Beryl orders the seven people gathered who had carried Rainbow Crystals, intending to combine them into one huge youma. Ami is determined to protect Urawa. Sailor Moon tries to heal Endymion.

Rei falls for Mamoru and persuades him to go on a date with her, whilst Nephrite targets the caretaker of a local park.