Michael Portillo reaches the Rocky Mountains of Colorado at Canon City, from where he heads out into the spectacular gorge of the Arkansas River aboard the historic narrow-gauge Royal Gorge Railroad. Along the way, he learns how controversy over the construction of the railroad caused bullets to fly. In Colorado Springs, Michael discovers an unexpected British outpost where he is enlisted to join the cricket team. At the foot of Pike's Peak, a choir sings an iconic American hymn, composed at the turn of the 20th century and inspired by the magnificent views to be seen from the summit. Michael heads 14,115 feet above sea level to see it for himself, aboard the Pike's Peak Cog Railway.

Michael Portillo is blown away by the skyscrapers in Illinois's Windy City, where he discovers how modern Chicago's skyline replaced a largely wooden city, destroyed in a great fire shortly before his Appleton's Guide was published. Training with the city's firefighters, Michael learns today's fire department respond to half a million emergency calls a year. Michael steps up to the plate with the Joliet Slammers, stars of the US national game of baseball, then sings for his supper at a quintessentially American restaurant bearing his name! Downtown at the Moody Church, Michael tracks down a pair of evangelists who toured Britain and the United States by rail at the time of his Guide.

Michael Portillo crosses the Atlantic once more to ride the railroads of North America with his faithful Appleton's Guide to the United States and Canada. Amid breathtaking scenery, he encounters magnificent beasts, joins intrepid explorers and witnesses unique customs on an awesome 1,500-mile journey to recapture the excitement and promise of the 19th-century American Frontier. In St Louis, Michael ascends America's monument to the Wild West, the astonishing Gateway Arch, the tallest free-standing monument in the United States, and makes a delightful discovery inside it. And on the banks of the Missouri River, he is invited aboard a magnificent replica of the original keel boat used for a historic expedition.

Following in the footsteps of European settlers, Michael Portillo rolls westwards across the United States. With true frontier spirit, he discovers the hidden pleasures of 19th-century railroad workers in Sedalia, known as the Sodom and Gomorrah of the West, and discovers the birthplace of ragtime and its most famous composer, Scott Joplin. Aboard a horse-drawn wagon in Independence, Michael confronts the brutal hardships faced by early pioneers on the wagon trail and discovers a living history museum town where the clock stopped in 1855. He ends this leg in the rail hub of Kansas City, Missouri, where freight trains can be a mile long.

On the next leg of his American adventure, Michael Portillo hits cowboy central in the former meat-packing capital of America - Kansas City, Missouri. In Paola, Kansas, Michael auctions livestock to the latter-day John Waynes of the state and then dines out on smoky spare ribs. In the West Bottoms district, he explores the city's Irish heritage and discovers the first Irish business opened in America, still family- owned and run by the great granddaughter of the founder. In St Joseph, Missouri, Michael discovers the legendary Pony Express, not a train but a mail service, and investigates the treacherous death of the outlaw Jesse James.

Michael Portillo is in Albuquerque, New Mexico, a city situated on the Rio Grande River. Once the headquarters of the Acheson, Topeka and Santa Fe Railroad, Albuquerque has treasures in store for rail fans, including an enormous locomotive weighing over 450 tonnes. It is being restored by volunteers, and Michael lends a hand. Michael then feels the heat of New Mexico's chilli pepper in a restaurant and learns how to make enchiladas. Michael joins the Grand Canyon Railway at Williams, Arizona, to reach one of the most spectacular sights on Earth. He learns how the 7-million-year-old gorge of the Colorado River was preserved for the nation.

Michael Portillo hits ranching territory in Las Animas County, Colorado, where he follows in the footsteps of 19th-century dudes from the east coast who came to experience life as cowboys. First up - how to use a lasso. At Bent's Old Fort, Michael learns about the Mexican-American War of 1848 and finds himself inspecting the troops. He also looks at America's relationship with firearms at a NRA shooting centre in Raton, New Mexico, and discovers how guns won the West. Back across the state line in Pueblo, Colorado, Michael is drawn to a giant steel mill mentioned in his Appleton's that produces rails a quarter of a mile long, transported on special trains.

Michael Portillo continues his 1,500-mile journey through the American Wild West, armed with his 19th-century Appleton's Guide. In Lawrence, Kansas, Michael enjoys a prairie chicken dance with a student of the Haskell Indian Nations University and learns how Native American Indians were treated in the 19th century. Michael joins the Jayhawks basketball team in the famous Allen Field house stadium with energetic encouragement from their cheerleaders, before travelling through Tornado Alley to Topeka to meet one of the first storm chasers in America. Driving out on the Great Plains, Michael learns about the Tallgrass Prairie and comes face to face with a herd of wild buffalo.

Michael Portillo continues his journey from the northern state of Minnesota to the Deep South. Michael discovers the story of Hiawatha and his lover Minnehaha, an epic poem featuring American Indian characters. He then meets a Dakota Sioux expert on Native American culture to learn about a dark chapter in United States history and how it is marked today. Michael has to put his diplomatic skills to the test at a Swedish-American lunch, where the centrepiece of the menu is reconstituted dried cod. And there is a taste of the golden age of luxury rail travel as Michael bounces on a bed in a beautifully restored Pullman carriage.

Michael Portillo's first stop aboard the Southwest Chief is Dodge City, Kansas, a famous frontier town of the Old West, where he visits the notorious Long Branch Saloon, scene of many shoot-outs. He finds the Dodge City Cowboy Band performing at the city depot and breaks bread with a descendant of a railroad land agent who sold thousands of acres of land to 19th-century European settlers. Near Lamar, Colorado, Michael visits the scene of a terrible massacre of American Indians, who found themselves in the way of white settlement of the Great Plains.

Armed with his 19th-century Appleton's Guide, Michael Portillo continues his 1,000-mile journey, beginning and ending on the Mississippi River from Minnesota to Tennessee. Riding the mainline of mid-America, Michael stops at rural Mattoon, where he gets a taste of the tough early life which shaped President Abraham Lincoln. Wiping the sweat from his brow, Michael struggles to split one rail compared with Lincoln's estimated 700 a day. Basket in hand, Michael joins the Schwartz family apple harvest in Centralia and learns how to make apple butter. He uncovers industrial unrest in the coal mines of Carbondale then heads to Kentucky and the banks of the Mississippi, where a bloody conflict unfolded which proved decisive in victory for Lincoln's Union.

Michael Portillo arrives in Santa Fe, the state capital of New Mexico, and once the capital of a Spanish kingdom. He explores the beautiful colonial architecture of the city and is invited to visit a Native American pueblo atop a 367-foot-high sandstone bluff. Michael eavesdrops on rehearsals for Puccini's Girl of the Golden West at the glorious Santa Fe Opera House, and at the Governor's Palace learns about the author of the biblical epic Ben Hur. Finally, Michael catches up with two of the famous Harvey Girls at the La Fonda Hotel to hear about Fred Harvey, the railroad caterer from Lancashire who made his fortune in America.

Michael Portillo continues his journey from the northern state of Minnesota to the Deep South. Steered by his Appleton's Guidebook, Michael travels along the Upper Mississippi to Red Wing, Minnesota, and the home of Laura Ingalls Wilder, author of Little House on the Prairie. Joining the Empire Builder rail service, he travels to Winona, where he finds out how in the 19th century the US Army Corps of Engineers made the Mississippi navigable. The river is maintained today not just for freight, but to host bald eagles and pelicans too. Michael joins in the dancing at a tribal gathering of the Dakota Sioux, and in La Crosse learns about the Native American origins of a fast-moving, combative sport.

Michael Portillo continues his journey from the northern state of Minnesota to the Deep South. Michael reaches Tomah, Wisconsin, where he helps harvest cranberries, an important crop for the US. Boarding the Empire Builder rail service once again, he travels to Wisconsin Dells, where an innovative photographer first captured motion. Michael takes a plunge in the waterpark capital of the world and finds a pleasant surprise in Baraboo - the circus is in town and it travels by train. Guided by a former clown and ringmaster, Michael explores the train's spectacular wagons.

Steered by his Appleton's Guide, Michael Portillo continues his 1,000-mile rail journey south from Chicago through Kankakee to Champaign, Illinois. In full swing on the fairway at the Flossmoor Country Club, Michael discovers how wealthy businessmen from the city flocked to play and how the railroads fostered the growth of suburban life. Beside the Kankakee River, Michael is invited to visit the Frank Lloyd Wright-designed house that changed the face of American architecture. On the platform at Kankakee station, Michael parties with the locals as they celebrate the City of New Orleans rail service, immortalised in song by Arlo Guthrie. He gets his hands on a vintage hooter riding on the Monticello Heritage rail line and in Champaign learns a thing or two at a railroad university.

Armed with his Appleton's Guide, Michael Portillo travels the lower Mississippi aboard a paddle steamer to hear about the life and work of former river boat captain, Samuel Clemens, better known as Mark Twain. In the city of Memphis, Tennessee, Michael visits the historic Elmwood Cemetery, where he uncovers the story of a devastating epidemic, which claimed the lives of thousands. He discovers one of the largest rail freight hubs in the United States and in the home of the blues, meets contemporary artist Cedric Burnside in studio before joining millions of Elvis fans at Graceland. An invitation to a duck palace and the honorary position of Duck Master carry curious responsibilities at the 19th-century Peabody Hotel.

Michael continues his American rail journey west from St Louis to Jefferson City, Missouri, following the tracks of European settlers in the 19th century. He begins this leg at the birthplace of a rural icon in Washington, Missouri, where he attempts to craft a corn cob pipe. In Hermann, Missouri, Michael tucks in to bratwurst as he discovers the descendants of German settlers in the town continue their traditions today. And in the Missouri state capital, Jefferson City, Michael finds an enormous fortified building that served as the jail for the entire Wild West.

Michael Portillo continues his 1,000-mile journey from the northern state of Minnesota to the home of the blues in Memphis, Tennessee. Today, in the nation's rail capital, where tracks pass underground and overground and are elevated into the air, Michael investigates the ultimate marshalling yard. At the ornate Palmer House Hotel, Michael teams up with the head chef to recreate the original chocolate brownie, invented by Bertha Palmer for Chicago's World Exposition in 1893. He discovers the origins of the Sanitary and Ship Canal and uncovers the history of an incredible civil engineering project which raised the city to new heights. Heading deep underground, Michael inspects a modern day scheme on a similarly awesome scale, described by the boss as the largest toilet in the world.

Michael Portillo begins a new journey from Minnesota's Twin Cities in the north of the US to Memphis, Tennessee, in the Deep South. In this leg, Michael discovers how Minneapolis, part of one metropolitan area with its immediate neighbour, St Paul, harnessed the power of the mighty Mississippi to become a great industrial centre. It is also the artificial limb capital of the world. In St Paul, meanwhile, Michael visits the birthplace of F Scott Fitzgerald and meets a jazz age trumpeter and Fitzgerald fan who introduces him to the Jay Gatsby lifestyle.

Michael Portillo continues his journey from the northern state of Minnesota to the Deep South. Michael is in Milwaukee, located on the shore of Lake Michigan and home of the Harley-Davidson. Michael learns about the history of motorcycles there, the first of which was made in 1903, and hitches a ride on the back of a modern version. German gymnastics is Michael's next challenge as he joins the Ladies' Auxiliary Exercise Class, a legacy of Milwaukee's 19th-century German settler community which is still going strong today. In Racine, Michael discovers a man who knew how to sort the wheat from the chaff and made a business out of it.