Michael Portillo continues his American rail journey through New England as he heads for the Canadian border. First stop is Burlington, Vermont, a busy timber port at the time of his Appleton's guide. Michael ventures deep into the forest to learn how sustainable and technological innovations have transformed the state's billion dollar logging industry. Following the old trade route across Lake Champlain, he hears of a pivotal battle during the War of 1812 where a British defeat gave the United States a new confidence on the world stage. In Plattsburg, Michael learns of the surprising origins of a classic Christmas carol. Lead by his guidebook, he travels into the wilderness of the Adirondack Mountains. Here the rich and famous of Appleton's day established great camps to get back to nature, in the lap of luxury. He visits the largest of the camps, reached by boat and even a private funicular railway. In Lake Placid, Michael braves the steep curves and speeds of an Olympic bobsleigh run. Last stop is an American fort mistakenly built in Canada!

Michael Portillo begins a new journey on the tracks of the Ocean line to explore Canada’s maritime provinces en route to Quebec City. Clutching his 1899 Appleton’s Guide to Canada, he begins in the Atlantic port of Halifax, Nova Scotia, where he discovers an 18th-century British hilltop citadel, manned at the time of his guide by the 78th Highland Regiment. Michael joins the men who recreate the roles of those Scottish soldiers today. At the mercy of the young 'sergeant major', Michael learns the drill in kilt and sporran. Michael follows his Appleton’s to a vast Victorian dry dock, still in use today by shipbuilders for the Royal Canadian Navy and finds out what it takes to build a state-of-the-art Arctic Patrol vessel. He learns of a catastrophic explosion in Halifax harbour in 1917, which killed 2,000 people and left 25,000 homeless. Former residents of an African-Canadian community torn in two by the railway tell Michael of their struggle for redress.

Clutching his 1899 copy of Appleton’s Guide to Canada, Michael Portillo travels on the Ocean train from Nova Scotia to New Brunswick. Along the way, he investigates the world’s biggest tide at Hopewell Rocks and admires its dramatic rock formations and caves. Michael apparently defies gravity on a magnetic hill in a 1965 Pontiac Bonneville. North of Moncton in Miramichi, he joins the Elsipogtog First Nation in a pow wow, where he learns about quilting and traditional dress. In Amherst, Michael investigates the history of an ambitious ship railway designed to ferry ships by rail over the isthmus between the Bay of Fundy and the Northumberland Strait. He quarries highly-prized Wallace sandstone for a 150-year-old family firm.

Michael Portillo explores the province of Quebec with his 19th-century Appleton’s Guide to Canada. He takes the fabulously scenic Charlevoix train along the north bank of the mighty St Lawrence River to La Malbaie. Following his guidebook to the beautiful basilica at Sainte-Anne-de-Beaupre, Michael discovers the racks of crutches discarded by the healed and meets modern-day visitors in search of miracles. The Train de Charlevoix, built to transport pilgrims, now conveys tourists along the north bank of the St Lawrence River to the Murray lakes. Michael tours the fine 19th-century houses, which were once the haunt of the Gatsby generation. Taking to the skies in a seaplane, Michael flies over the Laurentian Mountains to land on an isolated lake, where he fishes for trout for his supper.

Steered by his 1899 Appleton’s Guide, Michael Portillo strikes west across Manitoba into the province of Saskatchewan. High above the prairie at Riding Mountain, Michael discovers how a middle-class British boy from Hastings transformed himself into an influential indigenous naturalist called Grey Owl. Deep in the prairie, Michael finds a network of railways that once served the wheat farmers of Saskatchewan and learns how communities grew up around the grain elevators used to load the crop on to rail wagons. The Wheatland Express welcomes a new recruit to the sidings on the afternoon shift. At Manitou Beach, Michael reaches the Dead Sea of Canada, a 14-mile lake three times saltier than the ocean. A Yellow Quill First Nations elder tells Michael about the healing properties of the water, and Michael tries it for himself.

Michael Portillo continues west through the Canadian Prairie on his thousand-mile rail journey from Winnipeg, Manitoba, to Jasper, Alberta. Following his 1899 Appleton’s guide, Michael explores a glossy, glassy, oil-rich Edmonton, second city of Alberta. On the banks of the North Saskatchewan River, he travels three centuries back in time to experience the life of les voyageurs, who travelled huge distances within Canada by foot and canoe to trade fur with indigenous people. Michael admires Edmonton’s early 20th-century heritage streetcars, preserved by the Radial Railway Society, and seizes the chance to drive one across a spectacular high-level bridge over the North Saskatchewan River. Edmonton prides itself on its modern light rail system, offering rapid transit to 80 million passengers per year. Michael hears how this growing city plans to keep pace. His journey across Canada’s vast open spaces reaches a dramatic scenic conclusion in the Rocky Mountains.

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Michael Portillo's railway journey across eastern Canada concludes in the nation's largest metropolis, Toronto. He begins his Toronto tour at Union Station. Now busier than the city's international airport, Michael is shown the ambitious engineering works underground to support the growing number of commuters. From the dig down, he boldly goes to the dizzying heights at the CN Tower for an extreme outdoor experience at the top of the structure. Nerves are calmed at the Royal York Hotel, one of a network of luxury hotels built by the railway known as the 'castles of the north'. Catching the street car, Michael finds out how Toronto made itself a magnet for money after it set up its own stock exchange, but not before he presses the button to open the day's trading. Ending his time in the city's High Park, he seeks out the origins of a celebrated Canadian song that helped to shape the maple leaf as the country's national symbol.

In Baltimore, home of the first railroad in the United States, the Baltimore and Ohio, he discovers how the first American steam engine, the Tom Thumb, owed much to pioneering British technology. He investigates race relations in the troubled city, taking a drive downtown with a former drug dealer, now a teacher. On the city's beautiful east coast, Michael discovers the impressive star-shaped Fort McHenry and learns how the Star-Spangled Banner national anthem was born. Medics at the city's Johns Hopkins Hospital show Michael how their institution has grown from its 19th-century foundation by the railroad magnate into a world-leading centre for healthcare. And at the city's Lexington Market, Michael learns what gives a Maryland crab cake the edge.

Michael continues his American journey in Manhattan's Lower East Side, where he narrowly avoids a scrap with an historic gang of New York and visits the grim tenement buildings where thousands of the city's immigrants lived and worked. In the West Side, Michael discovers how a once lethal run of track has been transformed into a public park, raised above the city streets. Forsaking the rails for a ferry, Michael heads for Ellis Island, where some 12 million immigrants entered America. Michael is given a privileged tour of the gleaming new transport hub under construction close to the site of Ground Zero.

Using his 1899 Appleton's guide, Michael Portillo's rail odyssey through eastern Canada continues along the Grand Trunk railway, following the route of the St Lawrence River. At Brockville, he leaves the tracks for a nautical pilgrimage through the beautiful Thousand Islands. In the port city of Kingston, Ontario, Michael visits Fort Henry and, dressed for the occasion, is entrusted to fire the naval guns that protected the nation's southern border during the 19th century. Travelling west to Port Hope, he learns of the antics of a celebrated 19th-century high-wire walker known as The Great Farini. And, in the spirit of showmanship, Michael tests his balance with the modern sport of slack lining. This leg of the journey ends in Oshawa at the opulent home of the McLaughlin family, who helped build a new economy for Canada when they switched from manufacturing carriages to motor cars.

Michael Portillo's thousand-mile rail journey through California continues south toward Santa Clara in the heart of Silicon Valley and ends in the surf of Santa Cruz. In a high-tech bar, where robots wait on the customers, Michael traces the origins of modern-day search engines and social media to an institution founded by a railroad baron. At tech giant Intel, Michael glimpses the future on a test drive in an autonomous car. At San Jose, the biggest city in the valley, Michael visits Japantown to meet members of a far eastern community who settled here peacefully during the 19th century but faced hostility during the mid-20th. He is then beaten over the head and body by youngsters a third of his age as he attempts to learn the martial art of kendo. A ride aboard the Roaring Camp and Big Trees Railroad takes Michael through groves of mammoth trees - the magnificent California Redwoods. Emerging from the forest on a second vintage railroad, Michael hits the beach and like a true beach boy, he takes to the waves - on a surfboard.

Michael hitches a ride with the Amish in a horse-drawn buggy through rich countryside settled in the 18th century by religiously oppressed Europeans. Charmed by their modest way of life, he watches as they sell their beautiful quilts at auction. In Strasburg, known as Traintown USA, Michael joins the crew of the oldest continuously operated railroad in the United States. After oiling the magnificent engine, he rides on the footplate of the vintage steam locomotive. His last stop on this leg is Gettysburg, the most famous battlefield of the American Civil War, where, in 1863, President Abraham Lincoln made a momentous speech.

Michael reaches a milestone on his American journey: the boundary between the northern and southern states, known as the Mason-Dixon line. He discovers the origins of what became for black Americans the border between slavery and freedom, in an 18th-century English dispute over land. On the Wilmington and Western Railroad, Michael meets the passenger train Phoebe Snow, created by the railroads at the turn of the 20th century to reassure passengers that clean burning coal wouldn't make their clothes dirty. Michael uncovers the explosive history of gunpowder production in Delaware, begun by a Frenchman, whose chateau still stands. He then takes a boat trip up the Susquehanna River following the route taken by one of the first English settlers, John Smith.

Michael Portillo's 19th-century Appleton's guidebook leads him to the Parker House Hotel, where in his best pinny, he whisks up a Boston cream pie.

Led by his Appleton's guidebook and tracing the footsteps of the Pilgrim Fathers, Michael Portillo heads for Plymouth, the home town of America. He learns how indigenous tribes of Wamponoag people taught the newly arrived settlers to live off the land, the inspiration for one of the biggest holidays in the American calendar.