Now that the estate is finally in good shape, Archie feels he must do something with himself. So when Killwillie decides to open a new restaurant in New York, Archie jumps at the chance to manage it. Meanwhile, Lexie scrambles to save a local couple from being evicted and Duncan discovers a shattering secret about his family.

While repairs are being done in the cellar, the family is living in Duncan's B&B. Paul returns to Glenbogle, stirring unwanted feelings within Molly.

Duncan takes a dip into the dating pool with the help of Molly's new dating agency while Paul gets working on establishing his new climbing center. But when the locals find out Paul is targeting out of area teenagers, they refuse to help in developing the center.

Golly gets an unexpected surprise when his daughter Jess shows up and Duncan travels to Glasgow for a job interview. Meanwhile, Paul takes on new responsibilities by trying to reform the local football team.

Duncan starts his new job with Dorothy and Hermione, leaving the Head Ranger position open at Glenbogle. While Jess tries to impress Golly to secure the position, Paul hires a young delinquent with many hidden talents.

Archie's elder sister Lizzie turns up with a life changing proposition for him and Andrew has his own proposition with Molly. At the same time, Archie and Paul continue to battle over joint control of the estate.

Now that Archie has left, Lexie has taken control over all estate affairs leaving Paul feeling left out. Golly is convinced that Andrew cannot be trusted and confronts Molly about his inclination.

Hermione returns to Glenbogle and begins to stalk Duncan with dark intentions. Meanwhile, Dorothy seizes control over a small fishing island and gets Ewan fired from the local pub.

Lexie is the first female laird of Glenbogle and the host of the annual clan gathering which is sure to start creating troubles in every area of the estate. Matters go from bad to worse when an ancient rival of the MacDonalds' sets foot on enemy land.

Lexie gets an unexpected and unwelcome surprise when her father shows up at Glenbogle. Father-daughter relations are strained between Golly and Jess as well, and Paul learns of a secret that Amanda's been hiding.