Archie introduces a pack of wolves into Glenbogle even though Golly has warned him against it. The new cook/housekeeper arrives causing a rift between Archie and Lexie. Duncan has his work cut out when he must take a test to show he can look after the wolves when the Wildlife Centre opens. Archie swaps Lexie's ring pull engagement ring for a real diamond ring.

The day of the opening of the Glenbogle Wildlife Centre arrives, with nothing but trouble. A new employee, Esme, sabotages the signs and an enemy of Molly's decides wether the Centre will open or not. Meanwhile, Golly is fighting for Irene's affections with Duncan.

Two faces from the past turn up. Firstly, Katrina, and an old friend of Hector's, Louis. Neither of their intentions are honourable. Katrina wants Archie back and Louis wants control of Glenbogle by marrying Molly. Lexie is hurt by Katrina's prescence and packs a bag and goes over the hills, to be then tracked by Archie. Everyone arrives and the now happy couple get married.

David Fraser, an old friend of the MacDonald's, mainly Jamie's turns up with his freind Jonathon. Lexie takes a disliking to him, as he makes a pass at her. Archie thinks he is the same person as when they were younger, but only Lexie can see what he really is. She confronts Archie, who then has a fight with David, and he is thrown out of Glenbogle. Meanwhile Molly is harbouring a wolf runt (called Hector) so he won't be killed.

Duncan accidently booked a group of naturists onto the campsite, and when a wolf escapes, there is havoc all around. Duncan is also in trouble when Golly finds out about his relationship with Irene. When the wolf is recaptured Golly explains about his recent lapses and his bad eye sight. He then resigns from Glenbogle and leaves.

With Golly gone, Archie is lost without him. Lexie though locates him and gives him food and supplies. Golly comes out of hiding though when a school boy from a party goes missing and Golly finds him. Meanwhile Molly and Irene go to an art exhibition to impress on a dealer who has been to see her. Irene though spots someone she once knew, which starts to tell the truth of why she escaped to Glenbogle. Golly returns in the end when Archie finds some money for a procedure to save his eyesight.

Greg MacDonald and his wife challenge Archie for the role of Laird of Glenbogle. Also the debt from Hector's death duties must now be paid, £250,000. In order to save Glenbogle Molly agrees to marry Kilwillie, but then has a change of heart. They duel and Archie wins. Meanwhile, Irene tells Duncan she is pregnant and he cannot be the father.

Archie decides he has no work to do and so starts to decorate a room for the nursery. Lexie and Auntie Liz are trying to find the ghost of a late MacDonald. Andrew a sculptor arrives in Glenbogle as a new neighbour, and does not get off to a good start with Molly. The ghost is found out to be a private detective trying to find Irene sent by her jilted boyfriend. She leaves and Duncan is heartbroken

A stranger, Paul, turns up looking for Golly, believing Golly is his father. He is not Golly's son but is someone from Glenbogle's son. Archie and Duncan hunt for Big Eric because he is badly injured. Lexie is in charge of arranging the Ghillies's Ball, but things start to get on top of her, like people trying to help. Archie, reluctantly, lets Paul spend the night, and watches the ball from the side.

Molly now knows the truth about Paul and decides to leave with Andrew Booth. Archie does not want Paul around anymore and Lexie must try to smooth things out. Molly asks Golly why he never told her the truth about Hector and Paul's mother and Archie is feeling guilt over his older brothers,(Jamie),death. His feelings of guilt go away a little bit as he saves Paul from drowning in the current. Molly looks within herself and decides to stay and Archie and Lexie finally get their honeymoon,thanks to Kilwillie!

Paul launches his new idea of turning Glenbogle into a luxury retreat when a wealthy couple come to stay at the house. But when Golly has a back injury and Molly rushes to his aid, Paul finds himself short staffed and strapped for cash.

Paul's goddaughter Amy shows up, a relentless troublemaker, and begins to shake up Ewan and Zoe's relationship. Then, Tam Cooper, a recently released convicted poacher returns to Glenbogle, creating havoc for Paul and Golly.

Archie's wayward sister Lizzie turns up with her New Age boyfriend Gerald and it seems the pair have big plans to turn Glenbogle into a kind of hippy retreat. Archie and Gerald end up at loggerheads, however, when some of their New Age ideals seem to involve Hector rediscovering his lost youth and taking an ill advised swim in the lake.

Alex has decided that the estate is not being run with a firm enough hand and takes it upon himself to bring his military organization to the Glenbogle household but it isn't long before everyone becomes annoyed with Alex. Meanwhile, while Molly is away, Golly strikes up a new friendship with a local dance teacher named Meg.

Archie tries to juggle his life in London with his new role as Laird of Glenbogle. He and Lexie make a hectic journey down to London for a special Highland night in his restaurant and he dashes back north just in time to see Katrina beat Kilwillie in the local elections. Understandably Justine's patience is wearing very thin.

Duncan takes a dip into the dating pool with the help of Molly's new dating agency while Paul gets working on establishing his new climbing center. But when the locals find out Paul is targeting out of area teenagers, they refuse to help in developing the center.

Competition sparks between Paul and Chester when they both want to market Isabel's relish. To decide who will be the winner, a duel is called to see who is the better marksman of the two. Meanwhile Meg feels disheartened when Golly gets people into her dance class through favors and Jess gets set up on Rob Roy Radio.

Ewan is shocked when he finds his dad with another woman and learns that his parents are planning on separating. Meanwhile, Lucy's parents are coming over to meet Paul and begin to hint about the possibility of a future wedding.

Young restaurant owner Archie MacDonald is called urgently from London back to his ailing fathers bedside in the ancestral home in Scotland, Glenbogle. He soon discovers that Hector has nothing worse than a cold and his sweet but dotty mother Molly has called him home on a pretext and has worked events so he can't easily escape back to London.

While repairs are being done in the cellar, the family is living in Duncan's B&B. Paul returns to Glenbogle, stirring unwanted feelings within Molly.

Hermione returns to Glenbogle and begins to stalk Duncan with dark intentions. Meanwhile, Dorothy seizes control over a small fishing island and gets Ewan fired from the local pub.

Ever since Meg passed away, Golly has been taking care of baby Cameron as a single parent, with the help of his daughter, Jess. Meanwhile, as spring returns to Glenbogle, the list of repairs to be done is endless and Paul finds himself face to face with a feisty young shepherdess named Iona.

Golly gets an unexpected surprise when his daughter Jess shows up and Duncan travels to Glasgow for a job interview. Meanwhile, Paul takes on new responsibilities by trying to reform the local football team.

Duncan starts his new job with Dorothy and Hermione, leaving the Head Ranger position open at Glenbogle. While Jess tries to impress Golly to secure the position, Paul hires a young delinquent with many hidden talents.

Archie's elder sister Lizzie turns up with a life changing proposition for him and Andrew has his own proposition with Molly. At the same time, Archie and Paul continue to battle over joint control of the estate.