Scarlett’s breakdown is driving everyone to point fingers at who might be responsible for her condition. Deacon is blaming Rayna for pushing her too fast, Rayna is blaming Juliette and for forcing her on stage and Beverly thinks Rayna is only interested in seeing her label succeed. It always feels like everyone has an axe to grind with Juliette so she turns to Avery, only to find him preoccupied with Scarlett. During her moment of weakness, she is driven right into the hands of her nemesis. Meanwhile, Layla tries to convince Will to join her in a reality show.

Rayna resorts to drastic measures when she learns that Maddie has crossed paths with a duplicitous figure from her past. Juliette must choose how to publicly reckon with the truth of Jeff Fordham’s death—a decision that directly affects Layla. Scarlett and Gunnar consider parting ways as bandmates, and Will finally decides to do what he should have done long ago—step up as the leader of a cause and a community.

Rayna joins Scarlett on the road for her first tour date with Luke Wheeler and is blindsided when Deacon confronts her about putting too much pressure on Scarlett too soon. Jeff goes to war against Rayna on control of the album she produced with Liam, and Juliette makes a decision about Charlie Wentworth. Gunnar struggles to write a song for a major artist, and gets help from Scarlett-sparking an idea for Rayna that could potentially be liberating for her and life-changing for Gunnar.

Scarlett's career is on fire as she sings "Free" in concert with Zac Brown, but the demands of fame are burning her out. Rayna asks Deacon to help write a much-needed hit for her new album, and it's like old times--perhaps too much. Concert venues are dropping from Juliette's tour and Jeff, her label head, is furious, and Zoey is bruised by Gunnar's inattention when he gets an invitation to co-write from Jay DeMarcus of Rascal Flatts.

Rayna makes a last-ditch effort to protect her oldest daughter from the perils that befall young artists. Juliette reunites with her “Shenandoah Girl” co-star Noah West on the Oscar campaign trail, where she meets Kesha, who guest stars as herself. Elsewhere, Scarlett grows wary of Gunnar’s budding friendship with Autumn Chase; Layla fears falling into obscurity before her career even takes off; and Will suffers a devastating loss.

After his collapse while performing at the late night show, Will recovers in the hospital and frets about his future. Brad gets a restraining order against Deacon, fueling conflict between Deacon and Jessie. The media get wind of Juliette’s extended stay in South America and begin harassing Avery about it. Maddie gets caught up in the fast lane lifestyle of Jonah Ford.

Rayna and Teddy's relationship is tested when Rayna performs with Deacon at Teddy's campaign fundraiser and a woman from Teddy's past (Kimberly Williams-Paisley) shows up with a secret. Meanwhile, Juliette tries to burnish her image with an appearance on "Good Morning America" and is interviewed by Robin Roberts; Gunnar makes a connection with his publisher's assistant, Hailey; and Teddy and Coleman agree to take the moral high ground during the campaign -- but will it last?

Rayna thinks about moving on when Deacon sets boundaries in their relationship; as Juliette struggles with her mother's addictions, Deacon is reminded of difficult moments in his past; tension between Gunnar and Avery puts pressure on Scarlet.

After the crash, Rayna is in a coma and Deacon is left to wait on his pending jail sentence. Teddy is still shell-shocked about Peggy's pregnancy and struggles to understand where he belongs... Is it by Rayna's side with his daughters? Meanwhile, Maddie grapples with the revelation that Deacon is her father and, feeling alone, turns to Juliette, unwittingly revealing the details of what happened before the traumatic crash. And Gunnar and Scarlett's relationship becomes even more complicated after the proposal, so they turn to their friends, Will and Zoey, for support.

Juliette hits a career high, including singing a duet with Steven Tyler, but her personal life plummets; Avery takes care of baby Cadence at his parents' home in Texas; Rayna deals with the aftermath of Deacon's liver transplant.

A second aneurysm leaves Beverly brain dead and Deacon consumed with guilt, while Scarlett is forced to decide whether or not to keep Beverly on life support. Meanwhile, Juliette continues a destructive pattern of erratic behavior, pushing Avery to make the heart-wrenching decision to file for divorce.

Rayna wants to turn down Sony’s contract offer for Maddie to be a solo artist and finds her family life distracting her from her work with Markus. Luke is desperate to regain Colt’s trust; Gunnar’s girlfriend, Erin, causes issues on tour with Scarlett and Avery scrambles to make money doing insipid jingles.

Daphne struggles with the idea of Deacon dating. Juliette introduces Avery to Darius. Maddie and Jonah record a song, but Maddie finds Jonah’s life a little too public. Will learns news about Zach and find ways to enhance his appearance.

Darius helps Juliette unlock a dark truth about her past; Daphne and Jake bond over each other's dislike of their parents dating; Scarlett volunteers at an equine therapy ranch; Gunnar, Will and Avery struggle to find their sound.

Juliette discovers that Darius Enright’s Movement is not the happy utopia that it seemed to be. She demands to leave, but they won’t let her. The tension in the group too much to bear.

A visitor from the past comes into Deacon's life -- his father, a very unpleasant surprise. With Alannah out of the picture, the band flounders and the guys have to consider whether it's worth continuing.

Rayna learns her family is facing financial ruin; Juliette asks Deacon to write and tour with her exclusively; Juliette's mother re-enters her life.

Katie Couric guest stars as herself when Rayna appears on the "Katie" show, trying to set the record straight about her marriage and balancing her career with motherhood. Juliette attempts to take her album launch into her own hands but it backfires when she tweets out a surprise concert location. Maddie lies to Teddy about her whereabouts — with dangerous consequences — and Deacon is drawn to his dog’s pretty new veterinarian, Stacey. Meanwhile Avery tries to control his musical destiny by flouting Dominic and Marilyn, and things are awkward between Scarlett and Gunnar, as he continues to process his brother’s death.

Rayna and Deacon struggle to write their album. Maddie and Daphne create a disturbance at home. Damien apologizes to Scarlett. Scarlett and Gunnar question their relationship.

Maddie and Clay get pulled over by a police officer. Juliette searches for a hit to relaunch her career. Damien George returns to Nashville with a grand plan for Scarlett.

After a public meltdown, Juliette seeks to change her life from an unlikely source. Scarlett and Maddie take a trip. Maddie meets Jonah Ford, a famous pop star. Scarlett struggles to find herself. Will coaches Gunnar. Deacon deals with loneliness.

Deacon goes on a date with a familiar face. Juliette dives into a self-help movement. Will and Gunnar invite Avery to perform with them. Scarlett is reminded how her career as had a positive influence on fans. Daphne reaches out to Jake.

Daphne and Jake perform at school. Deacon and Jessie run into Brad. Alannah’s stage presence grows, creating a division in the band. Gunnar questions his relationship with Alannah. Maddie joins Jonah in Miami, bonds with Twig. Avery visits Juliette.

As Brad's country music singing competition begins, he and a singing coach instantly recognize Daphne as a star in the making. Meanwhile, Avery's relationship with Alannah deepens.

In the premiere episode, "Pilot," Rayna learns that her record label wants her to tour as the opening act for newcomer Juliette Barnes, and when she refuses, the stage is set for the ultimate power struggle for popularity. Complicating matters, Lamar's drive for power results in backing Teddy in a run for Mayor of Nashville, against Rayna's wishes.