8.6/101,511 votes#1 - The Collapse of Nature
Season 4 Episode 1 - Aired 4/14/2016
After two months of respite, Sarah’s hard-won refuge in Iceland is shattered by a Neolution attack. Once again forced to flee, she realizes no matter how far her family runs it will never be far enough.
Director: John Fawcett
Writer: Graeme Manson
8.3/101,230 votes#2 - Transgressive Border Crossing
Season 4 Episode 2 - Aired 4/21/2016
After evading a Neolution attack in Iceland, Sarah returns home to track down an elusive new ally. As Cosima struggles with Delphine’s disappearance, Alison and Donnie host a pregnant and ever-hungry Helena. Meanwhile Felix, feeling like an outsider, embarks on his own journey of self-discovery.
Director: John Fawcett
Writer: Russ Cochrane
8.1/101,137 votes#3 - The Stigmata of Progress
Season 4 Episode 3 - Aired 4/28/2016
As Sarah reels from the shocking discovery of Neolution’s biotech, she’s desperate to find answers. Rachel receives a visit from her keeper and takes a fraught step forward towards recovery. Meanwhile, Helena is mistaken for “Mrs. Hendrix” when she answers a visit from suspicious visitors.
Director: Ken Girotti
Writer: Aubrey Nealon
8.1/101,129 votes#4 - From Instinct to Rational Control
Season 4 Episode 4 - Aired 5/5/2016
Sarah realizes her goals have aligned with her once enemy and forms a precarious alliance in hopes of getting to the head of Neolution. Meanwhile, on her end, Alison dispatches Felix and Donnie to infiltrate a Neolution fertility clinic that Beth investigated. Back at the lab, Cosima and Scott receive a revolting delivery from Alison. As Rachel grows closer to Charlotte, Susan Duncan leaves a devastating decision in her hands.
Director: Peter Stebbings
Writer: Alex Levine
8.3/101,169 votes#5 - Human Raw Material
Season 4 Episode 5 - Aired 5/12/2016
Sarah’s day off to reconnect with Kira quickly turns into a sleuthing mission aimed squarely at discrediting Felix’s new ties. Cosima steps up with Donnie to further investigate the Neolutionist fertility research center. While Donnie bumps into an all too familiar face, Cosima makes a shocking discovery that leads her into a dangerous encounter.
Director: David Wellington
Writer: Kate Miles Melville
9.2/101,882 votes#6 - The Scandal of Altruism
Season 4 Episode 6 - Aired 5/19/2016
When Cosima admits doubts about finding a cure in time to save her life, Sarah makes a lone decision that exposes Kendall to those she’s spent years hiding from. Meanwhile, Art and Felix team up to protect naïve clone, Krystal Godderitch, from herself.
Director: Grant Harvey
Writer: Chris Roberts
7.9/101,189 votes#7 - The Antisocialism of Sex
Season 4 Episode 7 - Aired 5/26/2016
Sarah regresses to old habits while struggling to cope with the consequences of her decisions —reconnecting with Beth on a dark night of the soul. Back at the lab, Cosima, haunted by vicious memories and news of Delphine, attempts a dangerous Hail Mary to find a cure. At the Hendrixes, Alison is more determined than ever to create normalcy for her kids. As she forges ahead with a birthday party, Donnie is caught in the Neolution crosshairs.
Director: David Frazee
Writer: Graeme Manson, Nikolijne Troubetzkoy
8.3/101,113 votes#8 - The Redesign of Natural Objects
Season 4 Episode 8 - Aired 6/2/2016
As Alison is terrified by Neolution threats made against Donnie, Sarah and Mrs. S question her loyalty to her sisters. While Sarah is distracted, Siobhan sneaks out to seek vengeance. Meanwhile, Cosima and Susan Duncan work together in a final attempt to discover a cure.
Director: Aaron Morton
Writer: Peter Mohan
8.5/101,154 votes#9 - The Mitigation of Competition
Season 4 Episode 9 - Aired 6/9/2016
With Susan Duncan deposed, Sarah finds herself in the same position as Rachel — under the heel of a new HBIC — causing the archenemies to come together for the greater good. Cosima and Susan butt heads on the ethical implications of human experiments. Meanwhile, when the Hendrixes' legal counsel, Adele, further investigates Donnie’s case, she begins to suspect Felix is keeping deep secrets. Helena’s arrival forces Felix to make a difficult decision.
Director: David Frazee
Writer: Alex Levine
8.6/101,320 votes#10 - From Dancing Mice to Psychopaths
Season 4 Episode 10 - Aired 6/16/2016
When communication with Cosima is cut off and Neolution appears to be within striking distance, Sarah's alliance with Rachel goes out the window. Sarah lures a high-level Neolutionist into a trap, and sets out to bring down Neolution once and for all.
Director: John Fawcett
Writer: Graeme Manson
The Best Episodes of Orphan Black Season 4
Every episode of Orphan Black Season 4 ranked from best to worst. Discover the Best Episodes of Orphan Black Season 4!
A streetwise hustler is pulled into a compelling conspiracy after witnessing the suicide of a girl who looks just like her.
Genres:DramaSci-Fi & Fantasy
Season 4 Ratings Summary
"The Collapse of Nature" is the best rated episode of "Orphan Black" season 4. It scored 8.6/10 based on 1511 votes. Directed by John Fawcett and written by Graeme Manson, it aired on 4/14/2016. This episode is rated 0.3 points higher than the second-best, "Transgressive Border Crossing".