As Wilhelmina prepares for her flamboyant wedding, she quickly realizes that her maid of honor, Victoria Beckham, is stealing all her thunder. Meanwhile, Betty is filled with guilt about her pact with Wilhelmina to keep the latter's affair a secret and Ignacio prepares for his big day as he is about to become an official U.S. citizen. However, when Betty moves in with Henry the father/daughter relationship is strained.

Betty finds herself in a dilemma between Henry and Gio when each makes a romantic offer to her. Meanwhile, a softball game between ELLE and Mode has rivals Wilhelmina and Naomi Campbell facing off in the field. Finally, Hilda makes quite a discovery about Coach Diaz, and a surprise visit from an orphaned French boy stirs things up at Mode.

Betty's life is turned into turmoil after the numerous upsetting events that have recently occurred. Despite all this, she refuses to accept the fact that Henry's leaving has deeply affected her. Meanwhile, Amanda has a hard time coping with the news she recently discovered about who her real mother is and Wilhelmina plans to use all the recent tragedies to her advantage.

A crisis caused by Wilhelmina has the team at Mode struggling to meet a deadline under extreme time constraints. Consequently, Alexis is forced to flirt with the printer for an extension, and Betty and Daniel try to free a teen star out of rehabilitation for a re-shoot.

Wilhelmina tries to becomes part of the Meade family by any means necessary. Amanda withholds evidence and Betty applies a deadly perfume made by Fey Sommers.

Betty and Henry secretly go on a date to see Wicked, but Daniel, thinking that her date is Gio, gives two tickets to Betty and Gio and Gio does not want to give up those tickets. Meanwhile, Daniel gets in a meeting with an advertiser, a woman who has an interest in younger men, namely Daniel.

Following the wedding fiasco, Wilhelmina wastes no time and plots her next move to conquer Mode. Meanwhile, Betty tries her best to regain Daniel's affections.

Henry and Betty go on a double date with Hilda and Gio. Wilhelmina tries to change her image as the queen of mean but fails miserably.

Betty finds her attempts to suppress her feelings for Gio useless when the two end up chaperoning Justin's school dance together. Meanwhile, Amanda agrees to do a reality show with Gene, and Hilda goes out of her way to catch the eye of Coach Diaz. Finally, Wilhelmina makes her return to MODE, much to the distaste of the Meade family.

Betty gets a little overwhelmed about Wilhelmina's brand new hunky bodyguard. Plus, Henry returns to pay for insurance, which has Betty concerned about their relationship. Because of the fatal crash, Alexis gets amnesia and Bradford uses it to his advantage. Hilda comes to terms with recent events. And Amanda gets a surprise inheritance from her late biological mother.

Renee becomes convinced that Betty is in love with Daniel and thus starts a rivalry between the two for his affections. In the meantime, Gina returns to Queens to flash her new life and rich husband before Hilda's face.

Even though Henry is kind of back in her life, Betty finds herself annoyed and yet intrigued by the new sandwich guy, Giovanni. Meanwhile, everyone at Mode prepare for the Meades' Black and White Ball. Wilhelmina tries to get her wedding back on track. Amanda prepares to make her first appearance as Fey Sommers' birth child and fugitive Claire Meade invites herself to Wilhelmina's wedding.

Betty tries internet dating and she gets asked out; Marc attracts the attention of a photographer that Marc doesn't see as attractive; Wilhelmina tries to prove to the skeptical Daniel and Alexis that she can be part of the Meade family; Hilda is unaware that Justin has become a changed teen when he starts hanging with the wrong crowd and skips school.

When Justin and his classmates visit Mode, Betty is bothered by the message the magazine is sending across to the kids and encourages Daniel to use ordinary women as models during fashion week. Meanwhile, Amanda visits a psychic for news about her biological father and Wilhelmina gets an unprecedented turnout in her latest plan for absolute power.

Betty gets into trouble in her writing class when she submits a pre-written MODE story written by a woman who survived a bear attack (hence the title). Alexis returns to work, still with memory loss and having no idea how to act like a girl. Daniel receives a threat from their top advertiser who is against transsexuals and wants Daniel to fire his own sister. Amanda, still trying to figure out who her real father is, learns a shocking secret about Wilhelmina's past.

Betty is assigned to interview a novelist. Amanda and Marc may have uncovered who Amanda's biological father is. Daniel unknowingly dates Wilhelmina's sister.

Betty plans to spend her 24th birthday accompanied by Henry, but Charlie's sudden appearance jeopardizes her plans. Meanwhile, Wilhelmina plots against Renee and Daniel's relationship by hinting at secrets from a hidden past in her sister's life.

Betty decides to patch things up with Charlie by throwing her an unforgettable baby shower. Meanwhile, Hilda decides to confront Justin's teacher after learning that he is failing gym class and Christina learns who the father of Wilhelmina's baby really is.