The episode focuses on Betty Suarez struggling to tell her friend and boss, Daniel Meade about her new job in London. At the same time, Betty's sister, Hilda tries to keep their father from finding out she is moving to Manhattan. Wilhelmina Slater recovers from being shot by Tyler and Amanda Tanen finally finds her father.

As Hilda's wedding approaches, Betty asks Daniel to be her guest, Amanda tries to set Marc up with soap opera star Spencer Cannon, Justin wonders whether he can invite Austin, Wilhelmina finds a way to get the Meade empire and Betty gets an unexpected offer in both her personal and professional lives.

Betty, Matt and Amanda's love triangle in the Bahamas reaches boiling point. Betty has trouble with a photo shoot when Penelope Graybridge steals the shoot from her enemy Wilhemina. Marc falls for super sexy Troy.

Ignacio prepares a romantic dinner for his Jean. Betty and Hilda take the Be-Shure instant pregnancy tests. Meanwhile, Hartley gets Denise Ludwig to fill in for Wilhemina. However Marc plans to bring Willie back. Claire with Amanda track down Tyler, the son she gave up for adoption.

As Wilhelmina prepares for her flamboyant wedding, she quickly realizes that her maid of honor, Victoria Beckham, is stealing all her thunder. Meanwhile, Betty is filled with guilt about her pact with Wilhelmina to keep the latter's affair a secret and Ignacio prepares for his big day as he is about to become an official U.S. citizen. However, when Betty moves in with Henry the father/daughter relationship is strained.

The sudden death of a Mode editor leaves Betty and Marc competing for the same job. Meanwhile, Matt lets his jealousy over Henry get the best of him.

Betty invites Hilda and Justin to stay for dinner after Justin's acting class audition, but they decline. Living alone in her, Betty realizes that she misses Matt and fears that if something happened to her, no one would know. Though Amanda and Marc live in the same building, they do not want to walk to work with her, increasing her loneliness. Meanwhile, Marc is feeling tension working for both Daniel and Wilhelmina, and sees a career opportunity when he hears the vacant Senior Fashion Editor position will be filled.

In the episode, Betty Suarez finally has her braces removed, but only after she enters a dream sequence in which she sees what her life would be like if she had had perfect teeth. At the same time, Justin Suarez tries to contact Austin, who he kissed in the previous episode and Daniel Meade secures the Million Dollar Bra for a MODE cover shoot. Meanwhile Amanda Tanen reveals that her relationship with Tyler is getting serious.

Betty finds herself in a dilemma between Henry and Gio when each makes a romantic offer to her. Meanwhile, a softball game between ELLE and Mode has rivals Wilhelmina and Naomi Campbell facing off in the field. Finally, Hilda makes quite a discovery about Coach Diaz, and a surprise visit from an orphaned French boy stirs things up at Mode.

Hilda has doubts over her relationship with Archie after seeing an old flame Bobby Talercio. Marc helps Justin to fit in high school by telling him to become friends with some mean girls. Elsewhere Daniel is lured into a trap by Bennett. Wilhemina tries to come up with money to help Nico out of trouble.

Wilhemina is livid when she lays eyes on drag queen impersonator - Wilhediva Hater. Betty and Daniel go through a rough patch as she wins an award for her blog. Daniel has the press hot on his heels. While Marc is pleased with himself when a famous director does a photo shoot with him.

Justin is enjoying his acting class with his eccentric teacher, Lena Korvinka. When Betty thinks there might be a problem with him and his friends, Austin and Lilly, she drops by the class to check up on him. Betty gets sidetracked when a playwright called Zachary is introduced to the class. Betty tells Hilda about Zachary and believes his speech about reinvention was aimed at her.

Daniel reconnects with Grace Chin, a girl he once stood up for a date in college, and who is now New York City's top lawyer. Meanwhile, Betty befriends Charlie, an out-of-towner she meets at the Mode cafeteria, whom she later discovers to be Henry's girlfriend.

Betty's life is turned into turmoil after the numerous upsetting events that have recently occurred. Despite all this, she refuses to accept the fact that Henry's leaving has deeply affected her. Meanwhile, Amanda has a hard time coping with the news she recently discovered about who her real mother is and Wilhelmina plans to use all the recent tragedies to her advantage.

A crisis caused by Wilhelmina has the team at Mode struggling to meet a deadline under extreme time constraints. Consequently, Alexis is forced to flirt with the printer for an extension, and Betty and Daniel try to free a teen star out of rehabilitation for a re-shoot.

Wilhelmina tries to becomes part of the Meade family by any means necessary. Amanda withholds evidence and Betty applies a deadly perfume made by Fey Sommers.

Wilhelmina, getting ready for her first issue as editor, offers Betty a job as her assistant. Meanwhile, Daniel learns he will have to undergo a custody battle if he wants to keep Daniel Jr. in the U.S. Later, Hilda's romance with Coach Diaz continues to heat up.

A tragic accident leaves the Mode staff stunned and Betty must find out what really happened when a detective comes looking for answers. Meanwhile, Ignacio learns about Hilda and Coach Diaz.

Betty and Marc team up together to analyse Mode's budget. Matt continues with his pursuit to win Betty while she deals with some startling news on Facebook by a former flame

Betty finds that everything is changing as she begins her new job as associate features editor, especially since her boss is her ex-boyfriend, Matt Hartley, who is still unhappy about their breakup. In addition, Betty's new officemate, Megan, openly resents her, while Marc is constantly undermining her because he felt he was more deserving of the promotion. Meanwhile, without Betty as his assistant, Daniel is adrift, still mourning the loss of his wife, Molly. Betty tries to help Megan out on an assignment and connects with a thoughtful insect jewelry designer, Olivia, while Justin starts high school and is victimized by bullies-but won't tell his mom, Hilda. Claire Meade is finding that her power struggle at the company has her sitting pretty - for the moment -- opposite Wilhemina, who lost her claim to the Meade throne last season. But Wili has perhaps an even bigger problem than job security - she is harboring a secret at home that is driving Marc to distraction.

Betty gets to run her own Mode photo shoot at the United Nations to promote their anti-malaria initiative, "Nothing but Nets," but her big opportunity becomes a baptism by fire when Daniel's neediness takes its toll, and Matt openly demoralizes her. Meanwhile, Amanda does some soul-searching when she bonds with Daniel's new temp, Helen, a career receptionist as fabulous as Amanda is. Wili makes it her personal mission to protect a loved one, and Hilda reaches out to Marc to figure out how to help Justin.

Hilda is planning her wedding, which is getting closer and asks Betty about the plans for her bachelorette party. Betty tells her that she's planned a champagne tour of The Met museum, but Hilda and Bobby believe it's a joke and Betty has to think of something else. Bobby is put out when Justin turns down a lift to school in favour of walking with Austin.

Betty has always dreamed of working in the publishing world, but it seems as though there's no place for the non-thin and non-beautiful in the business. People already have their minds set in the idea that beautiful women are only the thin and perfect! Betty really wants a job in that wonderful, yet cruel, world. But can she really find a place in it?

Betty loses a book, which is the complete new edition of the magazine and is containing un-retouched photos of a popular actress. She has to find the book before the photos get out to the rest of the press. Meanwhile, Wilhelmina plots to use the snafu to bring Daniel down and Bradford Meade secretively inquires about Fey Sommers' final effects.

After Mode is forced to abandon its big Christmas photo shoot because another magazine has copied the idea, Betty forms an uneasy alliance with Marc and Amanda. The three fear they may have leaked the idea to someone at the rival mag. Meanwhile, Daniel and Wilhelmina reluctantly work together to come up with a new photo spread as Mode's deadline looms, and Betty's nephew, Justin, is oblivious to the turmoil as he spends a day at the magazine for a class project.