Sabo, the Chief of Staff of the Revolutionary Army, contacts Koala and everyone in Kamabakka Queendom. He explains to them about the things that he is accused of, but someone who was eavesdropping on the call takes a stunning action.

Luffy and Chopper fall into the rough cold sea along with the mysterious girl. Jimbei tries to help them but another attack comes from the unknown creature. Meanwhile, the Sunny is hit by torpedoes and the rest of the Straw Hats are thrown into the sea, too.

A young girl who came out of the big robot names herself Dr. Vegapunk and it surprises Franky and the rest of the Straw Hats. Meanwhile, Luffy’s group gets to the surface of the island where futuristic buildings stand and unknown giant creatures wander about.

While Luffy and the others are enjoying the futuristic machines of Egghead, someone who resembles Bartholomew Kuma comes after them. When Luffy tries to fight back, Bonney stops him and reveals her shocking relationship with Kuma.

The Blackbeard Pirates ambushed the Heart Pirates led by Law. The Blackbeard Pirates’ top officers use a variety of formidable Devil Fruit powers and keep pushing the Heart Pirates in order to obtain the copies of the Road Ponegliffs.

The Straw Hats without Luffy, Chopper, and Jimbei are taken to the laboratory on the clouds far above the ground by Lilith. They get excited about the things they've never seen before. However, inside the laboratory, a new threat is waiting for them.

Nami and the others are attacked by a Pacifista who resembles Jimbei and while they’re in battle, some of Vegapunk’s Satellites look on with excitement. Meanwhile, Luffy and the others find something extraordinary at the scrap heap.

Vegapunk Shaka shares with Robin and the others his theory about an ancient kingdom and the 100-Year Void. He even says that he visited Ohara after its destruction. Meanwhile, Luffy and Chopper try to activate the giant robot and something surprising happens.

Shaka reveals the secret behind the foundation of the Revolutionary Army and how the relationship between Vegapunk and Dragon affected it. Meanwhile, Vegapunk explains to Luffy and the others about his special power and how his clones work.

Vegapunk reveals the shocking history of the ancient giant robot and asks Luffy to take him off Egghead. Rob Lucci and his fellow Cipher Pol agents arrive at the island and ask for permission to enter the port.

CP-0 led by Rob Lucci makes a forced landing at Egghead and people on the Fabriophase are plunged into confusion while Shaka and the others prepare to fight back on the Labophase. The enraged Atlas stands up to defend the peace of the island.

Luffy transforms into the Warrior of Liberation as Rob Lucci also shows his awakened Devil Fruit form. As the two clash, Vegapunk reveals the shocking truth of the Gum-Gum Fruit and his eye-opening theory on Devil Fruits.

Vegapunk describes how he managed to grant the Seraphim Devil Fruit powers, which surprises Franky and the others. Meanwhile, Luffy is still facing Lucci in his savage, awakened form, but the rocket to the Labophase is about to leave.

Vegapunk asks the Straw Hats to take him on board their ship which makes Franky and Usopp happy but annoys Nami. Lucci orders his troops to block off all Vegapunk’s escape routes and looks to invade the heavily-secured Labophase.

CP-0 enters the Labophase and starts attacking the Thousand Sunny as soon as they find it. Luffy reunites with Nami and the others but finds out that Vegapunk is missing. Meanwhile, Bonney is pressing Vegapunk to change back her father Kuma.

The Seraphim come following CP-0 to the Labophase, pitting Zoro and Brook against the seven of them. Sanji, Franky, Edison, and Lilith leave the lab to fight them, but the Seraphim launch a full-scale attack on the lab.

Stussy suddenly knocked out Lucci to the Straw Hats’ surprise, and when Shaka reveals her true identity, they’re stunned again. Meanwhile, a major incident occurs on an island called Sphinx where Whitebeard came from.

CP-0 takes control of the Fabriophase but gets attacked by an unknown force. Meanwhile, in the Labophase, the Straw Hats and Dr. Vegapunk’s Satellites find out that Vegapunk has disappeared and begin searching for him.

Bonney witnesses her father Bartholomew Kuma’s painful memories. Meanwhile at the lab, Shaka notices that their communications and surveillance cameras are being cut off by somebody.

The Seraphim suddenly start attacking the Straw Hats and the Vegapunk Satellites. They are not stopping even when the Satellites order them to. When Luffy and Zoro are cornered, Lucci and Kaku wake up and make them a surprising offer.

Lucci and Kaku from CP-0, who came to Egghead with a mission to eliminate the seven Vegapunks, ask Luffy and Zoro to uncuff them so they can fight alongside them against the Seraphim, but Luffy and Zoro are reluctant to do so.

The Straw Hats and CP-0 relentlessly attack the Seraphim, but the Seraphim keep getting back up. Then, Zoro notices that the Seraphim remind him of someone who he fought before. Meanwhile, Shaka goes to check one place where he thinks Vegapunk Stella might be.

Stussy and Sentomaru discuss how big the threat that closes in on Egghead is as Luffy and the others keep fighting the uncontrollable Seraphim. Meanwhile, in an underground lab, someone approaches Vegapunk, who is being held captive.

Kid gives his crew a pep talk before the battle with Shanks. And Shanks also prepares for the battle but learns how his own subordinates do as they like to protect their beloved boss. Finally, Kid has his Damned Punk ready to shoot.

Perona breaks into the Blackbeard Pirates' prison to rescue her captain, Gecko Moria, and frees Koby, who then helps free other prisoners. Koby, now with a high bounty, distracts a mob of pirates to protect the others. He recalls Blackbeard's plan to trade him for Hachinosu's entry into the World Government. The pirates face attacks from clay golems, SWORD member Kujaku, and a Marine force led by Garp and Tashigi.